I have TreeLinesVisibility="Visible" on my TreeListView, as in the XAML snippet below.
<telerik:RadTreeListView x:Name="NEWCollectionControl" Grid.Row="1" EnableLostFocusSelectedState="False" ShowColumnHeaders="{Binding RelativeSource={RelativeSource Mode=FindAncestor, AncestorType={x:Type telerik:RadOutlookBar}}, Path=DataContext.Controller.ShowTreeHeader, Mode=TwoWay}" GridLinesVisibility="None" RowIndicatorVisibility="Collapsed" SelectionUnit="FullRow" ColumnWidth="*" TreeLinesVisibility="Visible" AutoExpandItems="{Binding RelativeSource={RelativeSource Mode=FindAncestor, AncestorType={x:Type telerik:RadOutlookBar}}, Path=DataContext.Controller.TreeViewAutoExpand, Mode=TwoWay}" AutoGenerateColumns="False" ItemsSource="{Binding Items}" IsExpandedBinding="{Binding IsExpanded, Mode=TwoWay}" SelectedCellsChanging="NEWCollectionControl_SelectedCellsChanging" DataLoaded="NEWCollectionControl_DataLoaded" Initialized="NEWCollectionControl_Initialized" SelectionChanged="NEWCollectionControl_SelectionChanged">
<telerik:RadTreeListView.ChildTableDefinitions>
<telerik:TreeListViewTableDefinition ItemsSource="{Binding Items}" />
</telerik:RadTreeListView.ChildTableDefinitions>
<telerik:RadTreeListView.Columns>
<telerik:GridViewImageColumn IsVisible="{Binding Controller.UseTreeViewIcons}" DataMemberBinding="{Binding Image}" TextAlignment="Left" ImageHeight="12" ImageWidth="{Binding ImageWidth}" />
<telerik:GridViewDataColumn DataMemberBinding="{Binding Translation.CurrentTranslation}" MinWidth="200" Header="{Binding Controller.PSRDictionary.Collection.CurrentTranslation}" />
</telerik:RadTreeListView.Columns>
</telerik:RadTreeListView>
However, the GridViewImageColumn, which was added later in order to allow for some icons to the side of the TreeListView, is causing elements of the 3rd hierarchy level onwards to not have those lines. The behaviour can be seen in the image below, and it will happen regardless if the column is visible or not.
I have tried to mess with many of the properties on these elements but to no success.